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Phylum
Taxonomic rank
Taxonomic rank
In biology, a phylum (; : phyla) is a level of classification, or taxonomic rank, that is below kingdom and above class. Traditionally, in botany the term division has been used instead of phylum, although the International Code of Nomenclature for algae, fungi, and plants accepts the terms as equivalent.{{cite book|title=The American Heritage New Dictionary of Cultural Literacy|chapter-url=http://dictionary.reference.com/browse/phylum|access-date=2008-10-04|edition=third|year=2005|publisher=Houghton Mifflin Company|chapter=Life sciences
General description
The term phylum was coined in 1866 by Ernst Haeckel from the Greek grc (φῦλον, "race, stock"), related to grc (φυλή, "tribe, clan"). Haeckel noted that species constantly evolved into new species that seemed to retain few consistent features among themselves and therefore few features that distinguished them as a group ("a self-contained unity"): "perhaps such a real and completely self-contained unity is the aggregate of all species which have gradually evolved from one and the same common original form, as, for example, all vertebrates. We name this aggregate [a] Stamm [i.e., stock / tribe] (Phylon)." In plant taxonomy, August W. Eichler (1883) classified plants into five groups named divisions, a term that remains in use today for groups of plants, algae and fungi. The definitions of zoological phyla have changed from their origins in the six Linnaean classes and the four embranchements of Georges Cuvier.
At its most basic, a phylum can be defined in two ways: as a group of organisms with a certain degree of morphological or developmental similarity (the phenetic definition), or a group of organisms with a certain degree of evolutionary relatedness (the phylogenetic definition). Attempting to define a level of the Linnean hierarchy without referring to (evolutionary) relatedness is unsatisfactory, but a phenetic definition is useful when addressing questions of a morphological nature—such as how successful different body plans were.
Definition based on genetic relation
The most important objective measure in the above definitions is the "certain degree" that defines how different organisms need to be members of different phyla. The minimal requirement is that all organisms in a phylum should be clearly more closely related to one another than to any other group. On the other hand, the highly parasitic phylum Mesozoa was divided into two phyla (Orthonectida and Rhombozoa) when it was discovered the Orthonectida are probably deuterostomes and the Rhombozoa protostomes.
This changeability of phyla has led some biologists to call for the concept of a phylum to be abandoned in favour of placing taxa in clades without any formal ranking of group size.
Definition based on body plan
A definition of a phylum based on body plan has been proposed by paleontologists Graham Budd and Sören Jensen (as Haeckel had done a century earlier). The definition was posited because extinct organisms are hardest to classify: they can be offshoots that diverged from a phylum's line before the characters that define the modern phylum were all acquired. By Budd and Jensen's definition, a phylum is defined by a set of characters shared by all its living representatives.
This approach brings some small problems—for instance, ancestral characters common to most members of a phylum may have been lost by some members. Also, this definition is based on an arbitrary point of time: the present. However, as it is character based, it is easy to apply to the fossil record. A greater problem is that it relies on a subjective decision about which groups of organisms should be considered as phyla.
The approach is useful because it makes it easy to classify extinct organisms as "stem groups" to the phyla with which they bear the most resemblance, based only on the taxonomically important similarities.
A classification using this definition may be strongly affected by the chance survival of rare groups, which can make a phylum much more diverse than it would be otherwise.
Known phyla
Animals
Main article: Animal
Total numbers are estimates; figures from different authors vary wildly, not least because some are based on described species, and some on extrapolations to numbers of undescribed species. For instance, around 25,000–27,000 species of nematodes have been described, while published estimates of the total number of nematode species include 10,000–20,000; 500,000; 10 million; and 100 million.
| Others |
|---|
| Phylum | Meaning | Common name | Distinguishing characteristic | Taxa described | ||||||||||||||||||||||
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| Agmata | Fragmented | Calcareous conical shells | 5 species, extinct | |||||||||||||||||||||||
| Annelida | last1 = Margulis | first1 = Lynn | author-link = Lynn Margulis | last2 = Chapman | title = Kingdoms and Domains: An Illustrated Guide to the Phyla of Life on Earth | publisher = Academic Press | edition = 4th corrected | date = 2009 | location = London | isbn = 978-0-12-373621-5 | url=https://books.google.com/books?id=9IWaqAOGyt4C}} | Segmented worms, annelids | Multiple circular segments | |||||||||||||
| Arthropoda | Jointed foot | Arthropods | Segmented bodies and jointed limbs, with chitin exoskeleton | + extant; 20,000+ extinct | ||||||||||||||||||||||
| Brachiopoda | 336}} | 336}} | Lophophore and pedicle | –500 extant; 12,000+ extinct | ||||||||||||||||||||||
| Bryozoa (Ectoprocta) | Moss animals | 332}} | Lophophore, no pedicle, ciliated tentacles, anus outside ring of cilia | extant | ||||||||||||||||||||||
| Chaetognatha | Longhair jaw | 342}} | Chitinous spines either side of head, fins | extant | ||||||||||||||||||||||
| Chordata | With a cord | Chordates | Hollow dorsal nerve cord, notochord, pharyngeal slits, endostyle, post-anal tail | + | ||||||||||||||||||||||
| Cnidaria | Stinging nettle | Cnidarians | Nematocysts (stinging cells) | |||||||||||||||||||||||
| Ctenophora | Comb bearer | 256}} | Eight "comb rows" of fused cilia | –150 extant | ||||||||||||||||||||||
| Cycliophora | Wheel carrying | Circular mouth surrounded by small cilia, sac-like bodies | + | |||||||||||||||||||||||
| Dicyemida | Lozenge animal | Single anteroposterior axial celled endoparasites, surrounded by ciliated cells | + | |||||||||||||||||||||||
| Echinodermata | Spiny skin | 348}} | Fivefold radial symmetry in living forms, mesodermal calcified spines | extant; approx. 13,000 extinct | ||||||||||||||||||||||
| Entoprocta | 292}} | Goblet worms | Anus inside ring of cilia | |||||||||||||||||||||||
| Gastrotricha | 288}} | Hairybellies | Two terminal adhesive tubes | |||||||||||||||||||||||
| Gnathostomulida | Jaw orifice | 260}} | Tiny worms related to rotifers with no body cavity | |||||||||||||||||||||||
| Hemichordata | 344}} | Acorn worms, hemichordates | Stomochord in collar, pharyngeal slits | extant | ||||||||||||||||||||||
| Kinorhyncha | Motion snout | Mud dragons | Eleven segments, each with a dorsal plate | |||||||||||||||||||||||
| Loricifera | Armour bearer | Brush heads | Umbrella-like scales at each end | |||||||||||||||||||||||
| Micrognathozoa | Tiny jaw animals | Accordion-like extensible thorax | ||||||||||||||||||||||||
| Mollusca | 320}} | Mollusks/molluscs | Muscular foot and mantle round shell | + extant; 80,000+ extinct | ||||||||||||||||||||||
| Monoblastozoa | ||||||||||||||||||||||||||
| (Nomen inquirendum) | One sprout animals | distinct anterior/posterior parts and being densely ciliated, especially around the "mouth" and "anus". | 1 | |||||||||||||||||||||||
| Nematoda | Thread like | 274}} | Round cross section, keratin cuticle | |||||||||||||||||||||||
| Nematomorpha | 276}} | 276}} | Long, thin parasitic worms closely related to nematodes | |||||||||||||||||||||||
| Nemertea | 270}} | 270}} | Unsegmented worms, with a proboscis housed in a cavity derived from the coelom called the rhynchocoel | |||||||||||||||||||||||
| Onychophora | Claw bearer | 328}} | Worm-like animal with legs tipped by chitinous claws | extant | ||||||||||||||||||||||
| Orthonectida | Straight swimmer | Parasitic, microscopic, simple, wormlike organisms | ||||||||||||||||||||||||
| Petalonamae | Shaped like leaves | An extinct phylum from the Ediacaran. They are bottom-dwelling and immobile, shaped like leaves (frondomorphs), feathers or spindles. | 3 classes, extinct | |||||||||||||||||||||||
| Phoronida | Zeus's mistress | Horseshoe worms | U-shaped gut | |||||||||||||||||||||||
| Placozoa | Plate animals | 242}} | Differentiated top and bottom surfaces, two ciliated cell layers, amoeboid fiber cells in between | + | ||||||||||||||||||||||
| Platyhelminthes | 262}} | 262}} | Flattened worms with no body cavity. Many are parasitic. | |||||||||||||||||||||||
| Porifera | Pore bearer | 246}} | Perforated interior wall, simplest of all known animals | extant | ||||||||||||||||||||||
| Priapulida | Little Priapus | Penis worms | Penis-shaped worms | |||||||||||||||||||||||
| Proarticulata | Before articulates | An extinct group of mattress-like organisms that display "glide symmetry." Found during the Ediacaran. | 3 classes, extinct | |||||||||||||||||||||||
| Rotifera | Wheel bearer | 282}} | Anterior crown of cilia | |||||||||||||||||||||||
| Saccorhytida | Saccus : "pocket" and "wrinkle" | Saccorhytus is only about 1 mm (1.3 mm) in size and is characterized by a spherical or hemispherical body with a prominent mouth. Its body is covered by a thick but flexible cuticle. It has a nodule above its mouth. Around its body are 8 openings in a truncated cone with radial folds. Considered to be a deuterostome or an early ecdysozoan. | 2 species, extinct | |||||||||||||||||||||||
| Tardigrada | Slow step | Water bears, moss piglets | Microscopic relatives of the arthropods, with a four segmented body and head | |||||||||||||||||||||||
| Trilobozoa | Three-lobed animal | Trilobozoans | A taxon of mostly discoidal organisms exhibiting tricentric symmetry. All are Ediacaran-aged | 18 genera, extinct | ||||||||||||||||||||||
| Vetulicolia | Ancient dweller | Vetulicolians | Might possibly be a subphylum of the chordates. Their body consists of two parts: a large front part and covered with a large "mouth" and a hundred round objects on each side that have been interpreted as gills or openings near the pharynx. Their posterior pharynx consists of 7 segments. | 15 species, extinct | ||||||||||||||||||||||
| Xenacoelomorpha | Strange hollow form | Xenacoelomorphs | last1=Cannon | first1=J.T. | last2=Vellutini | first2=B.C. | last3=Smith | first3=J. | last4=Ronquist | first4=F. | last5=Jondelius | first5=U. | last6=Hejnol | first6=A. | title=Xenacoelomorpha is the sister group to Nephrozoa | journal=Nature | volume=530 | issue=7588 | date=4 February 2016 | pages=89–93 | pmid=26842059 | doi=10.1038/nature16520 | url=http://urn.kb.se/resolve?urn=urn:nbn:se:nrm:diva-1844 | bibcode=2016Natur.530...89C | s2cid=205247296 }} | + |
| Total: 39 | 1,525,000 |
Plants
Main article: Plant
The kingdom Plantae is defined in various ways by different biologists (see Current definitions of Plantae). All definitions include the living embryophytes (land plants), to which may be added the two green algae divisions, Chlorophyta and Charophyta, to form the clade Viridiplantae. The table below follows the influential (though contentious) Cavalier-Smith system in equating "Plantae" with Archaeplastida, a group containing Viridiplantae and the algal Rhodophyta and Glaucophyta divisions.
The definition and classification of plants at the division level also varies from source to source, and has changed progressively in recent years. Thus some sources place horsetails in division Arthrophyta and ferns in division Monilophyta, while others place them both in Monilophyta, as shown below. The division Pinophyta may be used for all gymnosperms (i.e. including cycads, ginkgos and gnetophytes), or for conifers alone as below.
Since the first publication of the APG system in 1998, which proposed a classification of angiosperms up to the level of orders, many sources have preferred to treat ranks higher than orders as informal clades. Where formal ranks have been provided, the traditional divisions listed below have been reduced to a very much lower level, e.g. subclasses.
| Embryophyte (Land plants) |
|---|
| Division | Meaning | Common name | Distinguishing characteristics | Species described | ||||||||
|---|---|---|---|---|---|---|---|---|---|---|---|---|
| last = Mauseth | first = James D. | title = Botany: An Introduction to Plant Biology | edition = 5th | year = 2012 | isbn = 978-1-4496-6580-7 | publisher = Jones and Bartlett Learning | location = Sudbury, MA}} p. 489 | Anthoceros-like plants | Hornworts | Horn-shaped sporophytes, no vascular system | –300+ | |
| Bryophyta | Bryum-like plants, moss plants | Mosses | Persistent unbranched sporophytes, no vascular system | |||||||||
| Charophyta | Chara-like plants | Charophytes | ||||||||||
| Chlorophyta | 200}} | Chlorophytes | ||||||||||
| Cycadophyta | Cycas-like plants, palm-like plants | Cycads | Seeds, crown of compound leaves | –200 | ||||||||
| Ginkgophyta | Ginkgo-like plants | Ginkgophytes | Seeds not protected by fruit | extant; 50+ extinct | ||||||||
| Glaucophyta | Blue-green plants | Glaucophytes | ||||||||||
| Gnetophyta | Gnetum-like plants | Gnetophytes | Seeds and woody vascular system with vessels | |||||||||
| Lycophyta | Lycopodium-like plants | |||||||||||
| Clubmosses | Microphyll leaves, vascular system | extant | ||||||||||
| Angiospermae | Seed container | Flowering plants, angiosperms | Flowers and fruit, vascular system with vessels | |||||||||
| last1=Crandall-Stotler | first1=Barbara | last2=Stotler | first2=Raymond E. | year=2000 | chapter=Morphology and classification of the Marchantiophyta | page=21 | editor1=A. Jonathan Shaw | editor2=Bernard Goffinet | title=Bryophyte Biology | location=Cambridge | publisher=Cambridge University Press | isbn=978-0-521-66097-6 }} |
| Marchantia-like plants | ||||||||||||
| Liverworts | Ephemeral unbranched sporophytes, no vascular system | |||||||||||
| Polypodiophyta | Polypodium-like plants | |||||||||||
| Ferns | Megaphyll leaves, vascular system | |||||||||||
| Picozoa | Extremely small animals | Picozoans, picobiliphytes | 1 | |||||||||
| Pinophyta, | ||||||||||||
| Pinus-like plants | ||||||||||||
| Conifers | Cones containing seeds and wood composed of tracheids | extant | ||||||||||
| Prasinodermophyta | Prasinoderma-like plants | Picozoans, picobiliphytes, biliphytes | 8 | |||||||||
| Rhodophyta | Red plants | Red algae | Use phycobiliproteins as accessory pigments. | |||||||||
| Total: 16 |
Fungi
Main article: Fungi
| Division | Meaning | Common name | Distinguishing characteristics | Species described | ||
|---|---|---|---|---|---|---|
| Ascomycota | 396}} | 396}} sac fungi | Tend to have fruiting bodies (ascocarp). Filamentous, producing hyphae separated by septa. Can reproduce asexually. | 30,000 | ||
| Basidiomycota | 402}} | 402}} club fungi | title=Archaeal Genetics Boundless Microbiology | url=https://courses.lumenlearning.com/boundless-microbiology/chapter/archaeal-genetics/ | website=courses.lumenlearning.com}} | 31,515 |
| Blastocladiomycota | Offshoot branch fungus{{cite web | url = http://comenius.susqu.edu/biol/202/fungi/blastocladiomycota/default.htm | title = Blastocladiomycota | last1 = Holt | ||
| Chytridiomycota | Little cooking pot fungus{{cite web | url = http://comenius.susqu.edu/biol/202/fungi/chytridiomycota/default.htm | title = Chytridiomycota | last1 = Holt | ||
| Glomeromycota | 394}} | Glomeromycetes, AM fungi | Mainly arbuscular mycorrhizae present, terrestrial with a small presence on wetlands. Reproduction is asexual but requires plant roots. | 284 | ||
| Microsporidia | Small seeds{{cite web | url = http://comenius.susqu.edu/biol/202/fungi/microsporidia/default.htm | title = Microsporidia | last1 = Holt | ||
| Neocallimastigomycota | New beautiful whip fungus{{cite web | url = http://comenius.susqu.edu/biol/202/fungi/neocallimastigomycota/default.htm | title = Neocallimastigomycota | last1 = Holt | ||
| Zygomycota | 392}} | 392}} | Most are saprobes and reproduce sexually and asexually. | approx. 1060 | ||
| Total: 8 |
Phylum Microsporidia is generally included in kingdom Fungi, though its exact relations remain uncertain, and it is considered a protozoan by the International Society of Protistologists (see Protista, below). Molecular analysis of Zygomycota has found it to be polyphyletic (its members do not share an immediate ancestor),{{cite journal | display-authors = 3
Protists
Main article: Taxonomy of Protista
Kingdom Protista (or Protoctista) is included in the traditional five- or six-kingdom model, where it can be defined as containing all eukaryotes that are not plants, animals, or fungi. Protista is a paraphyletic taxon, which is less acceptable to present-day biologists than in the past. Proposals have been made to divide it among several new kingdoms, such as Protozoa and Chromista in the Cavalier-Smith system.
Protist taxonomy has long been unstable, with different approaches and definitions resulting in many competing classification schemes. Many of the phyla listed below are used by the Catalogue of Life, and correspond to the Protozoa-Chromista scheme, with updates from the latest (2022) publication by Cavalier-Smith. Other phyla are used commonly by other authors, and are adapted from the system used by the International Society of Protistologists (ISP). Some of the descriptions are based on the 2019 revision of eukaryotes by the ISP.
| Orphan groups |
|---|
| Phylum | Meaning | Common name | Distinguishing characteristics | Species described | Image | ||||||||||||||||||||||||||||||||||||||||||||||
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| Amoebozoa | Amorphous animals | Amoebozoans | Presence of pseudopodia for amoeboid movement, tubular cristae. | [[File:Amoeba_proteus.jpg | 100px]] | ||||||||||||||||||||||||||||||||||||||||||||||
| Apicomplexa | Apical infolds | Apicomplexans, sporozoans | Mostly parasitic, at least one stage of the life cycle with flattened subpellicular vesicles and a complete apical complex, non-photosynthetic apicoplast. | [[File:Toxoplasma gondii.jpg | 100px]] | ||||||||||||||||||||||||||||||||||||||||||||||
| Apusozoa(paraphyletic) | Apusomonas-like animals | Gliding biciliates with two or three connectors between centrioles | 32 | [[File:Podomonas_kaiyoae_C.jpg | 100px]] | ||||||||||||||||||||||||||||||||||||||||||||||
| Bigyra | Two rings | Stramenopiles with a double helix in ciliary transition zone | [[File:Aplanonet3.jpg | 100px]] | |||||||||||||||||||||||||||||||||||||||||||||||
| Cercozoa | Flagellated animal | Cercozoans | Defined by molecular phylogeny, lacking distinctive morphological or behavioural characters. | [[File:Euglypha_sp.jpg | 100px]] | ||||||||||||||||||||||||||||||||||||||||||||||
| Chromerida | Chromera-like organisms | Q30662251}} | Biflagellates, chloroplasts with four membranes, incomplete apical complex, cortical alveoli, tubular cristae. | 8 | [[File:Vitrella_brassicaformis_LM_Michalek_2020.png | 100px]] | |||||||||||||||||||||||||||||||||||||||||||||
| Choanozoa(paraphyletic) | Funnel animals | Opisthokont protists | Filose pseudopods; some with a colar of microvilli surrounding a flagellum | [[File:Desmarella_moniliformis.jpg | 100px]] | ||||||||||||||||||||||||||||||||||||||||||||||
| Ciliophora | Cilia bearers | Ciliates | Presence of multiple cilia and a cytostome. | [[File:Paramecium_bursaria.jpg | 100px]] | ||||||||||||||||||||||||||||||||||||||||||||||
| Cryptista | Hidden | Defined by molecular phylogeny, flat cristae. | [[File:Rhodomonas_salina_CCMP_322.jpg | 100px]] | |||||||||||||||||||||||||||||||||||||||||||||||
| Dinoflagellata | Whirling flagellates | Dinoflagellates | Biflagellates with a transverse ribbon-like flagellum with multiple waves beating to the cell's left and a longitudinal flagellum beating posteriorly with only one or few waves. | extant955 fossil | [[File:CSIRO ScienceImage 6736 dinoflagellate.jpg | 100px]] | |||||||||||||||||||||||||||||||||||||||||||||
| Endomyxa | Q28212529}} | Defined by molecular phylogeny, typically plasmodial endoparasites of other eukaryotes. | [[File:Vampyrella_lateritia.jpg | 100px]] | |||||||||||||||||||||||||||||||||||||||||||||||
| Eolouka(paraphyletic) | vauthors=Cavalier-Smith T | date=2013 | title=Early evolution of eukaryote feeding modes, cell structural diversity, and classification of the protozoan phyla Loukozoa, Sulcozoa, and Choanozoa | journal=European Journal of Protistology | volume=49 | issue=2 | pages=115–178 | doi=10.1016/j.ejop.2012.06.001 | pmid=23085100}} | Heterotrophic biflagellates with ventral feeding groove. | [[File:Stygiella incarcerata.jpg | 100px]] | |||||||||||||||||||||||||||||||||||||||
| Euglenozoa | True eye animals | Biflagellates, one of the two cilia inserted into an apical or subapical pocket, unique ciliary configuration. | extant20 fossil | [[File:Euglenoid_movement.jpg | 100px]] | ||||||||||||||||||||||||||||||||||||||||||||||
| Haptista | Fasten | Thin microtubule-based appendages for feeding (haptonema in haptophytes, axopodia in centrohelids), complex mineralized scales. | extant1,205 fossil | [[File:Coccolithus_pelagicus_2.jpg | 100px]] | ||||||||||||||||||||||||||||||||||||||||||||||
| Hemimastigophora | Incomplete or atypical flagellates | Hemimastigotes | Q85570914}} | [[File:Hemimastix_amphikineta.png | 60px]] | ||||||||||||||||||||||||||||||||||||||||||||||
| last=Pánek | first=Tomáš | last2=Tice | first2=Alexander K. | last3=Corre | first3=Pia | last4=Hrubá | first4=Pavla | last5=Žihala | first5=David | last6=Kamikawa | first6=Ryoma | last7=Yazaki | first7=Euki | last8=Shiratori | first8=Takashi | last9=Kume | first9=Keitaro | last10=Hashimoto | first10=Tetsuo | last11=Ishida | first11=Ken-ichiro | last12=Hradilová | first12=Miluše | last13=Silberman | first13=Jeffrey D. | last14=Roger | first14=Andrew | last15=Inagaki | first15=Yuji | last16=Eliáš | first16=Marek | last17=Brown | first17=Matthew W. | last18=Čepička | first18=Ivan | title=An expanded phylogenomic analysis of Heterolobosea reveals the deep relationships, non-canonical genetic codes, and cryptic flagellate stages in the group | journal=Molecular Phylogenetics and Evolution | volume=204 | date=16 January 2025 | doi=10.1016/j.ympev.2025.108289 | article-number=108289 | pmid=39826589 | url=https://linkinghub.elsevier.com/retrieve/pii/S1055-7903(25)00006-5 | url-access=subscription | doi-access=free}}Percolozoa | Percolomonas-like animals | Heteroloboseans, amoebomastigotes | Complex life cycle containing amoebae, flagellates and cysts. Amoeboflagellates with an amoeba, a flagellate, and a cyst stage in their life cycles. Amoebae usually cylindrical, with a monopodial locomotive form, relatively fast-moving via eruptive lobopodia. Flagellates usually with two or four flagella that arise at the anterior end of a feeding groove. Golgi apparatus lacking a classic stacked form. Mitochondria with discoidal cristae, some species with acristate, hydrogen-producing mitochondrion-related organelles. | [[File:Naegleria fowleri lifecycle stages.JPG | 100px]] | |
| Malawimonada | Malawimonas-like organisms | Malawimonads | Small free-living bicilates with two kinetosomes, one or two vanes in posterior cilium. | [[File:Malawimonasms.jpg | 100px]] | ||||||||||||||||||||||||||||||||||||||||||||||
| Metamonada | Middle monads | Metamonads | Anaerobic or microaerophilic, some without mitochondria; four kinetosomes per kinetid | [[File:Giardia muris trophozoite SEM 11643.jpg | 100px]] | ||||||||||||||||||||||||||||||||||||||||||||||
| Ochrophyta;Heterokontophyta | Ochre plants, heterokont plants | Heterokont algae, stramenochromes, ochrophytes, heterokontophytes | Biflagellates with tripartite mastigonemes, chloroplasts with four membranes and chlorophylls a and c, tubular cristae. | extant2,262 fossil | [[File:Diatoms_through_the_microscope.jpg | 100px]] | |||||||||||||||||||||||||||||||||||||||||||||
| Opisthosporidia | |||||||||||||||||||||||||||||||||||||||||||||||||||
| (often considered fungi) | last1=Karpov | first1=Sergey | last2=Mamkaeva | first2=Maria A. | last3=Aleoshin | first3=Vladimir | last4=Nassonova | first4=Elena | last5=Lilje | first5=Osu | last6=Gleason | first6=Frank H. | date=2014-01-01 | title=Morphology, phylogeny, and ecology of the aphelids (Aphelidea, Opisthokonta) and proposal for the new superphylum Opisthosporidia | journal= Frontiers in Microbiology | volume=5 | page=112 | doi=10.3389/fmicb.2014.00112 | pmc=3975115 | pmid=24734027 | doi-access=free}} | Parasites with chitinous spores and extrusive host-invasion apparatus | [[File:Fibrillanosema_spore.jpg | 100px]] | |||||||||||||||||||||||||||
| Perkinsozoa | Perkinsus-like animals | Perkinsozoans, perkinsids | Parasitic biflagellates, incomplete apical complex, formation of zoosporangia or undifferentiated cells via a hypha-like tube. | [[File:Dinovorax_pyriformis_PMC5609580_fig1c.png | 100px]] | ||||||||||||||||||||||||||||||||||||||||||||||
| Provora | Devouring voracious protists | Q115933632}} | [[File:Outline_drawing_of_Ubysseya_fretuma.svg | 50px]] | |||||||||||||||||||||||||||||||||||||||||||||||
| Pseudofungi | False fungi | Q28303534}} | [[File:Phytophthora_cactorum.jpg | 100px]] | |||||||||||||||||||||||||||||||||||||||||||||||
| Retaria | Reticulopodia-bearing organisms | Q28261633}} | extant50,000 fossil | [[File:Foraminifères_de_Ngapali.jpg | 100px]] | ||||||||||||||||||||||||||||||||||||||||||||||
| Sulcozoa(paraphyletic) | Groove-bearing animals | Aerobic flagellates (none, 1, 2 or 4 flagella) with dorsal semi-rigid pellicle of one or two submembrane dense layers, ventral feeding groove, branching ventral pseudopodia, typically filose. | + | [[File:Collodictyon_telophase_constrict.jpg | 70px]] | ||||||||||||||||||||||||||||||||||||||||||||||
| Telonemia | Telonema-like organisms | Telonemids | last1=Shalchian-Tabrizi | first1=K | last2=Eikrem | first2=W | last3=Klaveness | first3=D | last4=Vaulot | first4=D | last5=Minge | first5=M.A | last6=Le Gall | first6=F | last7=Romari | first7=K | last8=Throndsen | first8=J | last9=Botnen | first9=A | last10=Massana | first10=R | last11=Thomsen | first11=H.A | last12=Jakobsen | first12=K.S | title=Telonemia, a new protist phylum with affinity to chromist lineages | journal=Proceedings of the Royal Society B: Biological Sciences | date=28 April 2006 | volume=273 | issue=1595 | pages=1833–1842 | doi=10.1098/rspb.2006.3515 | pmid=16790418 | pmc=1634789 }} | [[File:Telonema_rivulare_(electron_micrography).jpg | 100px]] | ||||||||||||||
| Total: 26, but see below. |
The number of protist phyla varies greatly from one classification to the next. The Catalogue of Life includes Rhodophyta and Glaucophyta in kingdom Plantae, but other systems consider these phyla part of Protista. In addition, less popular classification schemes unite Ochrophyta and Pseudofungi under one phylum, Gyrista, and all alveolates except ciliates in one phylum Myzozoa, later lowered in rank and included in a paraphyletic phylum Miozoa. Even within a phylum, other phylum-level ranks appear, such as the case of Bacillariophyta (diatoms) within Ochrophyta. These differences became irrelevant after the adoption of a cladistic approach by the ISP, where taxonomic ranks are excluded from the classifications after being considered superfluous and unstable. Many authors prefer this usage, which lead to the Chromista-Protozoa scheme becoming obsolete.
Bacteria
Main article: Bacterial phyla
Currently there are 41 bacterial phyla (not including "Cyanobacteria") that have been validly published according to the Bacteriological Code
- Abditibacteriota
- Acidobacteriota, phenotypically diverse and mostly uncultured
- Actinomycetota, High-G+C Gram positive species
- Aquificota, deep-branching
- Armatimonadota
- Atribacterota
- Bacillota, Low-G+C Gram positive species, such as the spore-formers Bacilli (aerobic) and Clostridia (anaerobic)
- Bacteroidota
- Balneolota
- Bdellovibrionota
- Caldisericota, formerly candidate division OP5, Caldisericum exile is the sole representative
- Calditrichota
- Campylobacterota
- Chlamydiota
- Chlorobiota, green sulphur bacteria
- Chloroflexota, green non-sulphur bacteria
- Chrysiogenota, only 3 genera (Chrysiogenes arsenatis, Desulfurispira natronophila, Desulfurispirillum alkaliphilum)
- Coprothermobacterota
- Deferribacterota
- Deinococcota, Deinococcus radiodurans and Thermus aquaticus are "commonly known" species of this phyla
- Dictyoglomota
- Elusimicrobiota, formerly candidate division Thermite Group 1
- Fibrobacterota
- Fusobacteriota
- Gemmatimonadota
- Ignavibacteriota
- Kiritimatiellota
- Lentisphaerota, formerly clade VadinBE97
- Mycoplasmatota, notable genus: Mycoplasma
- Myxococcota
- Nitrospinota
- Nitrospirota
- Planctomycetota
- Pseudomonadota, the most well-known phylum, containing species such as Escherichia coli or Pseudomonas aeruginosa
- Rhodothermota
- Spirochaetota, species include Borrelia burgdorferi, which causes Lyme disease
- Synergistota
- Thermodesulfobacteriota
- Thermomicrobiota
- Thermotogota, deep-branching
- Verrucomicrobiota
Archaea
Main article: Archaea
Currently there are 2 phyla that have been validly published according to the Bacteriological Code
- Nitrososphaerota
- Thermoproteota, second most common archaeal phylum Other phyla that have been proposed, but not validly named, include:
- "Euryarchaeota", most common archaeal phylum
- "Korarchaeota"
- "Nanoarchaeota", ultra-small symbiotes, single known species
Notes
References
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