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Gentoo penguin

Species of bird


Species of bird

The gentoo penguin ( ) (Pygoscelis papua) is a penguin species (or possibly a species complex) in the genus Pygoscelis, most closely related to the Adélie penguin (P. adeliae) and the chinstrap penguin (P. antarcticus). The earliest scientific description was made in 1781 by Johann Reinhold Forster with a type locality in the Falkland Islands. The species calls in a variety of ways, but the most frequently heard is a loud, trumpeting call, emitted with the head thrown back.

Names

The application of "gentoo" to the penguin is unclear. Gentoo was an Anglo-Indian term to distinguish Hindus from Muslims. The English term may have originated from the Portuguese gentio ("pagan, gentile"). Some speculate that the white patch on the bird's head was thought to resemble a turban.

It may also be a variation of another name for this bird, "Johnny penguin", with Johnny being the English counterpart of the Spanish nickname Juanito and sounding vaguely like gentoo. The Johnny rook, a predator, is likely named after the Johnny penguin.

The specific name papua is a misnomer; in the original description, Johann Reinhold Forster, a naturalist who had circumnavigated the world with Captain James Cook, mistakenly assumed that the species occurred in Papua (New Guinea), the closest gentoos being over 6000 km to the south (on Macquarie Island). No penguins are found in New Guinea. Others trace the error to a "possibly fraudulent claim" in 1776 by French naturalist Pierre Sonnerat, who also alleged a Papuan location for the king penguin despite never having been to the island himself.

Taxonomy

The gentoo penguin is one of three species in the genus Pygoscelis. Mitochondrial and nuclear DNA evidence suggests the genus split from other penguins around 38 million years ago (Mya), about 2 million years after the ancestors of the genus Aptenodytes. In turn, the Adélie penguins split from the other members of the genus around 19 Mya, and the chinstrap and gentoo penguins finally diverged around 14 Mya.

There are four subspecies recognized by the International Ornithological Congress:

ImageSubspeciesDistribution
[[File:2020-11 Kerguelen Islands - Gentoo penguin 11.jpg120px]]P. p. taeniata (eastern gentoo penguin) (Peale, 1849)Crozet Islands, Prince Edward Islands, Kerguelen Islands, Heard Island, and Macquarie Island
[[File:Falkland Islands Penguins 05.jpg120px]]P. p. papua (northern gentoo penguin) (Forster, 1781)Falkland Islands, Martillo Island in the Beagle Channel, and Isla de los Estados (Argentina)
[[File:Gentoo Penguin Baby (24940372635).jpg120px]]P. p. ellsworthi (southern gentoo penguin) Murphy, 1947the Antarctic Peninsula, the South Orkney Islands, South Shetland Islands and South Sandwich Islands
[[File:Gentoo Penguin at Cooper Bay, South Georgia.jpg120px]]P. p. poncetii (South Georgia gentoo penguin) Tyler, Bonfitto, Clucas, Reddy & Younger, 2020South Georgia Island

Although the population on the Kerguelen Islands is tentatively included in taeniata, it may also be a distinct subspecies.

Before 2021, only two subspecies of the gentoo penguin had been recognised: P. p. papua (subantarctic gentoo) and the smaller P. p. ellsworthi (Antarctic gentoo). However, a 2020 study suggested that the gentoo penguin be split into a species complex of four morphologically similar but separate species: the northern gentoo penguin (P. papua sensu stricto), the southern gentoo penguin (P. ellsworthi), the eastern gentoo penguin (P. taeniata) and the newly described South Georgia gentoo penguin (P. poncetii). The International Ornithological Congress incorporated the results of this study in 2021, but relegated the newly recognized or newly described species to subspecies of P. papua.

Description

The gentoo penguin is easily recognised by the broad, white stripe extending like a bonnet across the top of its head and its bright orange-red bill. It has pale whitish-pink, webbed feet and a relatively long tail – the most prominent tail of all penguin species. Chicks have grey backs with white fronts. As the gentoo penguin waddles along on land, its tail sticks out behind, sweeping from side to side, hence the scientific name Pygoscelis, which means "rump-tailed".

A close-up of head on the [[West Falkland

Gentoo penguins can reach a length of 70 to, making them the third-largest species of penguin after the emperor penguin and the king penguin. Males have a maximum weight around 8.5 kg just before moulting and a minimum weight of about 4.9 kg just before mating. For females, the maximum weight is 8.2 kg just before moulting, but their weight drops to as little as 4.5 kg when guarding the chicks in the nest. Birds from the north are on average 700 g heavier and 10 cm longer than the southern birds. Southern gentoo penguins reach 75 – in length. They are the fastest underwater swimmers of all penguins, reaching speeds up to 36 km/h. Gentoos are well adapted to frigid and harsh climates.

Breeding

Egg, [[Museum Wiesbaden

The breeding colonies of gentoo penguins are located on ice-free surfaces. Settlements can be located directly on the shoreline or considerably inland. They prefer shallow coastal areas and often nest between tufts of grass. In South Georgia, for example, breeding colonies are 2 km inland. In colonies farther inland, where the penguins nest in grassy areas, they shift their location slightly each year because the grass becomes trampled over time.

Gentoo penguins breed on many subantarctic islands. The main colonies are on the Falkland Islands, South Georgia and the South Sandwich Islands and Kerguelen Islands; smaller colonies are found on: Macquarie Island, Heard Islands, Crozet Islands, South Shetland Islands and the Antarctic Peninsula. The total breeding population is estimated to be over 600,000 birds. Nests are usually made from a roughly circular pile of stones and can be quite large, 20 cm high and 25 cm in diameter. The stones are jealously guarded, and their ownership can spark noisy disputes and physical attacks among individuals. They are also prized by the females, even to the point that a male penguin can obtain the favours of a female by offering her a choice stone.

Two eggs are laid, both weighing around 130 g. The parents share incubation and change duty daily. The eggs hatch after 34 to 36 days. The chicks remain in the nests for around 30 days before joining other chicks in the colony and forming crèches. The chicks moult into subadult plumage and go out to sea at around 80 to 100 days.

Diet

Gentoos mainly live on fish, crustaceans such as krill and shrimp, and cephalopods. In February and March, crustaceans make up about 10% of the diet, while from March to June, it is about 75%. From June to October, grey rockcod (Lepidonotothen squamifrons) make up 90% of their diet. Cephalopods make up only 10% of the diet throughout the year.

Gentoos are opportunistic feeders, and around the Falklands are known to take roughly equal proportions of fish (Patagonotothen sp., Thysanopsetta naresi, Micromesistius australis), squat lobsters (Munida gregaria) and squid (Loligo gahi, Gonatus antarcticus, Moroteuthis ingens). Other prey include Channichthys rhinoceratus and octopuses.

Physiology

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Gentoo penguins do not store as much fat as Adélie penguins, their closest relative; gentoos require less energy investment when hunting because the net gain of energy after hunting is greater in gentoos than Adélies. As embryos, gentoos require a lot of energy to develop. Oxygen consumption is high for a developing gentoo embryo. As the embryo grows and needs more oxygen, consumption increases exponentially until the gentoo chick hatches. By then, the chick is consuming around 1800 ml O2 per day.

Predators

South Georgia

In the sea, leopard seals, sea lions, and killer whales are all predators of the gentoo. On land, there are no predators of full-grown, healthy gentoo penguins. Skuas and giant petrels regularly kill many chicks and steal eggs; petrels kill injured and sick adult gentoos. Various other seabirds, such as the kelp gull and snowy sheathbill, also snatch chicks and eggs. Skuas on King George Island have been observed attacking and injuring adult gentoo penguins in apparent territorial disputes.

Conservation status

The population of Pygoscelis papua in the maritime Antarctic is rapidly increasing. Due to regional climate change, they colonise previously inaccessible territories to the south. , the IUCN Red List lists the gentoo as least concerned with a stable population trend. However, rapid declines in some key areas are believed to be driving a moderate overall decline in the species population. Examples include Bird Island, South Georgia, where the population has fallen by two-thirds over 25 years. Many threats to this species, including pollution, hunting, fishing, and human recreational activities, continue to affect them.

References

References

  1. BirdLife International. (2020). "''Pygoscelis papua''".
  2. Woods, R.W. (1975) ''Birds of the Falkland Islands'', Antony Nelson, Shropshire, UK.
  3. (1988). "Portuguese vocables in Asiatic languages: from the Portuguese original of Monsignor Sebastião Rodolfo Dalgado, Volume 1". Asian Educational Services.
  4. Chester, Jonathan. (1996). "Penguins: Birds of Distinction". Penguin Books.
  5. "gentoo penguin".
  6. Beaglehole, J.C.. (2017). "The Journals of Captain James Cook on his Voyages of Discovery". Taylor & Francis.
  7. Gorman, James. (1990). "The Total Penguin". Prentice Hall.
  8. Campbell, David G.. (2002). "The Crystal Desert: Summers in Antarctica". Houghton Mifflin Harcourt.
  9. Troelstra, Anne S.. (2017). "Bibliography of Natural History Travel Narratives". Brill.
  10. (2013). "Australian Bird Names". Csiro.
  11. (2006). "Multiple gene evidence for the expansion of extant penguins out of Antarctica due to global cooling". Proc Biol Sci.
  12. "Kagu, Sunbittern, tropicbirds, loons, penguins – IOC World Bird List".
  13. "Subspecies Updates – IOC World Bird List".
  14. environment.gov.au. "''Pygoscelis papua papua'' — Gentoo Penguin (subantarctic)".
  15. itis.gov. "''Pygoscelis papua'' (J. R. Forster, 1781)".
  16. (2020). "Morphometric and genetic evidence for four species of gentoo penguin". Ecology and Evolution.
  17. (2020). "Cryptic speciation in gentoo penguins is driven by geographic isolation and regional marine conditions: Unforeseen vulnerabilities to global change". Diversity and Distributions.
  18. "Gentoo penguin videos, photos and facts – ''Pygoscelis papua''".
  19. "''Pygoscelis papua'' (gentoo penguin)".
  20. "Gentoo Penguin – Pygoscelis Papua".
  21. "Gentoo penguin videos, photos and facts – ''Pygoscelis papua''".
  22. "Antarctica fact file wildlife, gentoo penguins".
  23. "Gentoo penguin". BBC Nature.
  24. (2011). "Pygoscelis papua (gentoo penguin)".
  25. Schmidt-Nielsen, K.. (1960). "The Salt-Secreting Gland of Marine Birds". Circulation.
  26. (2016). "Physiological differences between two overlapped breeding Antarctic penguins in a global change perspective". Polar Biology.
  27. Actams, N. J.. (1992). "Embryonic metabolism, energy budgets and cost of production of the king ''Aptenodytes patagonicus'' and gentoo ''Pygoscelis papua'' penguin eggs". Comparative Biochemistry and Physiology Part A: Physiology.
  28. February 2014 observation and photo report by Robert Runyard, translator for INACH (Chilean Antarctic Institute).
  29. International), BirdLife International (BirdLife. (2019-08-28). "IUCN Red List of Threatened Species: ''Pygoscelis papua''".
  30. (2021-11-17). "Gentoo Penguins (''Pygoscelis papua'') started using mosses as nesting material in the southernmost colony on the Antarctic Peninsula (Cape Tuxen, Graham Land)". Polar Biology.
  31. Negus, Christopher. (5 May 2008). "Linux Bible: Boot Up to Ubuntu, Fedora, KNOPPIX, Debian, openSUSE, and 11 Other Distributions". John Wiley & Sons.
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