Euornithes

Description

Clarke et al. (2006) found that the most primitive known euornithians (the Yanornithiformes) had a mosaic of advanced and primitive features. These species retained primitive features like gastralia and a pubic symphysis. They also showed the first fully modern pygostyles, and the type specimen of Yixianornis (IVPP 13631) preserves eight elongated rectrices (tail feathers) in a modern arrangement. No earlier pygostylians are known which preserve a fan of tail feathers of this sort; instead, they showed only paired plumes or a tuft of short feathers.

Classification

The name Euornithes has been used for a wide variety of avialan groups since it was first named by Leonhard Stejneger in 1884. It was first defined as a clade in 1998 by Paul Sereno, who made it the group of all animals closer to birds than to Enantiornithes (represented by Sinornis). This definition currently includes similar content as another widely used name, Ornithuromorpha, named and defined by Luis Chiappe in 1999 as the common ancestor of Patagopteryx, Vorona, and Ornithurae, plus all of its descendants. Because one definition is node-based and the other branch-based, Ornithuromorpha is a slightly less inclusive group.

Relationships

The cladogram below follows the results of a phylogenetic analysis by Lee et al., 2014:

The following cladogram below follows the results of a phylogenetic analysis by Pei et al., 2020:

Other genera

The following is a list of primitive euornithian genera and those that cannot be confidently referred to any subgroups, following Holtz (2011) unless otherwise noted.

• †Alamitornis
• †Changmaornis
• †Changzuiornis
• †Dingavis
• †Gargantuavis
• †Horezmavis
• †Iteravis
• †Juehuaornis
• †Platanavis
• †Wyleyia?
• †Yumenornis
• Xinghaiornis
• †Zhyraornis

Note that Holtz also included the genera Eurolimnornis and Piksi as euornitheans, though they have since been re-identified as pterosaurs.

References

References

1. (2015). "The oldest record of Ornithuromorpha from the Early Cretaceous of China". Nature Communications.
2. (2022). "Forty new specimens of ''Ichthyornis'' provide unprecedented insight into the postcranial morphology of crownward stem group birds". PeerJ.
3. (2006). "Insight into the evolution of avian flight from a new clade of Early Cretaceous ornithurines from China and the morphology of ''Yixianornis grabaui''". Journal of Anatomy.
4. Stejneger, Leonhard Hess. (1884). "Classification of birds". The Illustrated Science Monthly.
5. (2013). "Morphological Clocks in Paleontology, and a Mid-Cretaceous Origin of Crown Aves". Systematic Biology.
6. Rui Pei, Michael Pittman, Pablo A. Goloboff, Mark A. Norell, Stephen L. Brusatte, Xing Xu. (2020). "Potential for Powered Flight Neared by Most Close Avialan Relatives, but Few Crossed Its Thresholds". Current Biology.
7. Holtz, Thomas R. Jr. (2012) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,'' [http://www.geol.umd.edu/~tholtz/dinoappendix/HoltzappendixWinter2011.pdf Winter 2011 Appendix.]
8. (2013). "Previously Unrecognized Ornithuromorph Bird Diversity in the Early Cretaceous Changma Basin, Gansu Province, Northwestern China". PLOS ONE.
9. Huang, J., Wang, X., Hu, Y., Liu, J., Peteya, J. A., & Clarke, J. A. (2016). A new ornithurine from the Early Cretaceous of China sheds light on the evolution of early ecological and cranial diversity in birds. ''PeerJ'', '''4''': e1765. {{doi. 10.7717/peerj.1765
10. (2012). "Systematic reinterpretation of ''Piksi barbarulna'' Varricchio, 2002 from the Two Medicine Formation (Upper Cretaceous) of Western USA (Montana) as a pterosaur rather than a bird". Geodiversitas.