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Chorrillo Formation

Geologic formation in Patagonia, Argentina

Chorrillo Formation

Summary

Geologic formation in Patagonia, Argentina

FieldValue
nameChorrillo Formation
imagePerito Moreno ISS.jpg
captionSatellite image of the Chorrillo Formation site (in the middle on the right). In the bottom left is the Perito Moreno Glacier.
typeGeological formation
periodMaastrichtian
age
prilithologyClaystone
namedbyArbe & Hechem
year_ts1984
locationPatagonia
regionSanta Cruz Province
countryArgentina
coordinates
paleocoordinates
underliesCalafate Formation
overliesLa Irene Formation
thickness50 m
extentSoutheast of Viedma Lake
North of Argentino Lake
Austral Basin
map{{Location map+Argentina
relief1
width250
floatcenter
lat_deg-49.9
lon_deg-72.5
marksize12

North of Argentino Lake Austral Basin

The Chorrillo Formation, also named as Chorillo Formation, is a Maastrichtian (Late Cretaceous Epoch, 72.1–66 million years ago) geologic formation in southern Patagonia, Argentina. The formation is more than 50 m thick and underlies the Calafate Formation and rests on top of the La Irene Formation. It constitutes a significant uppermost Cretaceous continental sedimentary sequence within the Magallanes Basin, located in Santa Cruz Province, Argentina, approximately 30 km southwest of El Calafate. Initial investigations in the 1990s laid the groundwork, with detailed paleontological studies emerging since 2019, revealing a rich biotic assemblage. The formation correlates with the lower Dorotea Formation in Chile, sharing dinosaurian ecotypes, while both are clearly different from other northern Patagonian Maastrichtian units (e.g., Allen Formation, Los Alamitos Formation), which are dominated by saltasaurine sauropods and abelisaurids, suggesting potential temporal or environmental distinctions.

Geology

The Chorrillo Formation is embedded within the foreland Magallanes Basin, shaped by Late Cretaceous Andean orogeny driven by subduction along the Pacific margin. Tectonic subsidence, coupled with sediment supply from the rising Patagonian Andes, facilitated deposition. Lithologically, the unit comprises fine- to coarse-grained clastics: mudstones and siltstones dominate floodplain facies, often organic-rich and pedogenically modified; sandstones, arkosic to lithic, form channel fills with cross-bedding; and conglomerates appear as basal lags. Clay mineralogy is dominated by smectite, with minor illite and kaolinite, reflecting volcanic ash alteration. Sedimentologically, it records a meandering fluvial system on a distal floodplain, featuring fining-upward channel sequences, crevasse splays, and overbank deposits. High-frequency paleosol stacking indicates local hydrologic and topographic variability, while low-frequency patterns suggest avulsion-driven progradation. Regional ties extend to the Dorotea Formation in Chile, reflecting basin-wide tectonic influences.

Paleoenvironment

Modern equivalent, Valdivian Rainforest

Paleoenvironmental reconstructions indicate a temperate-warm, seasonally humid climate with mean annual temperatures of 10-18 °C and precipitation of 900–1500 mm, inferred from paleosols, clay mineralogy, and biotic indicators. Depositional settings encompass low-energy fluvial floodplains with paludal and lacustrine elements, marked by poor drainage, hydromorphic paleosols (e.g., vertisols, histosols), and redoximorphic features such as mottling and iron nodules, suggesting fluctuating water tables and episodic anoxia. Aquatic communities feature Nymphaeaceae, Salviniaceae, Salviniaceae, and Zygnemataceae, indicative of freshwater ponds and swamps, while terrestrial elements include conifers, ferns, and angiosperms in riparian zones. Floral phytozones, derived from palynological data, comprise five assemblages across the section, exhibiting quantitative variations in taxa proportions without major turnover: lower zones are fern-dominated (e.g., Cyatheaceae), transitioning to angiosperm-rich upper zones (e.g., Proteaceae, Nothofagidites), attributed to subtle humidity fluctuations or habitat gradients relative to fluvial channels. These patterns align with Maastrichtian assemblages in southern Gondwanan basins (e.g., La Anita, Lefipán Formation), reflecting angiosperm diversification and Weddellian Province expansion.

Fossils

Dinosaur remains are among the fossils that have been recovered from the formation.

Fossils recovered from the formation are:

Arthropoda

Decapoda

GenusSpeciesLocalityMaterialNotesImages
AstacideaIndeterminate"Locality 4"MPM 21516, three gastrolithsPreviously identified as crushing fish teeth[[File:Crayfish.jpgcenterthumb196x196pxExtant example]]

Insects

GenusSpeciesLocalityMaterialNotesImages
last1=Verafirst1=Ezequiel I.last2=Monferranfirst2=Mateo D.last3=Massaferrofirst3=Julietalast4=Sabaterfirst4=Lara M.last5=Gallegofirst5=Oscar F.last6=Perez Loinazefirst6=Valeria S.last7=Moyano-Pazfirst7=Damiánlast8=Agnolínfirst8=Federico L.last9=Manabefirst9=Makotolast10=Tsuhijifirst10=Takanobulast11=Novasfirst11=Fernando E.date=2023-12-11title=A Maastrichtian insect assemblage from Patagonia sheds light on arthropod diversity previous to the K/Pg eventjournal=Communications Biologyvolume=6issue=1doi=10.1038/s42003-023-05596-2issn=2399-3642pmc=10711029pmid=38072954article-number=1249}}IndeterminateLa Anita farmMPM-Pal 21835-3:W34/1; MPM-Pal 21835-23:W55/0; MPM-Pal 21835-22:R52/2; MPM-Pal 21835-20:Z47/3; MPM-Pal 21835-22: G55/0; MPM-Pal 2183-21: E42/3A Glossatan Lepidopteran
Diamesinae"Morphotype 3"La Anita farmMPM-Pal 21835-9:E34/4; MPM-Pal 21835-19: V34/4A Chironomidae Nematoceran. Similar to the living genus Paraheptagyia[[File:Potthastia.gaedii.-.lindsey.jpgcenterthumb196x196pxExtant example]]
EphemeropteraIndeterminateLa Anita farmMPM-Pal 21835-9:W37/1A Mayfly
LepidopteraIndeterminateLa Anita farmMPM-Pal 21835-10:X30/2; MPM-Pal 21835-7:S27/2
Orthocladiinae"Morphotype 1"La Anita farmMPM-Pal 21835-24: M31/0; MPM-Pal 21835-3: P46/4A Chironomidae Nematoceran
"Morphotype 2"La Anita farmMPM-Pal 21835-25: V26/3
"Morphotype 5"La Anita farmMPM-Pal 21835-9: Q58/3It resembles modern Cricotopus[[File:Midges.jpgcenterthumb196x196px[[Cricotopus]]]]
Tanypodinae"Morphotype 4"La Anita farmMPM-Pal 21835-2:Y40/3A Chironomidae Nematoceran[[File:Tanypodinae - probably a melanic Procladius species - Flickr - gailhampshire.jpgcenterthumb196x196pxExtant example]]

Molluscs

Gastropods

GenusSpeciesLocalityMaterialNotesImages
last=Miquelfirst=S. E.year=2024title=New species and records of freshwater and land gastropods from the Cretaceous of Argentinajournal=Neues Jahrbuch für Geologie und Paläontologie - Abhandlungenvolume=310issue=2pages=119–131doi=10.1127/njgpa/2023/1173}}A. agnolini "Locality 4"ShellsAn achatinid gastropod[[File:Allopeas gracile 003.jpgcenterthumb196x196pxExtant specimen]]
BocourtiaB. leonardodavincii"Locality 4"ShellsA bulimulid gastropod[[File:Scutalus tupacii (MNHN-IM-2000-24710).jpegcenterthumb187x187pxExtant specimen]]
CoelostemmaC. patagonica"Locality 4"ShellsA holospirid gastropod
HolospiraH. sp."Locality 4"ShellsAn urocoptidae gastropod
LeptinariaCf.L. sp."Locality 4"ShellsAn achatinid gastropod[[File:Obese Funnel (Leptinaria unilamellata).jpgcenterthumb237x237pxExtant specimen]]
LilloiconchaL. novasi"Locality 4"ShellsA charopid gastropod
NaesiotusCf.N. sp."Locality 4"ShellsA bulimulid gastropod[[File:Naesiotus stenogyroides.pngcenterthumb178x178pxExtant specimen]]
PhysaP. sp."Locality 4"ShellsA physidae gastropod[[File:Physa Marmorata, shell.jpgcenterthumb178x178pxExtant specimen]]
PhysellaCf.P. sp."Locality 4"ShellsA physidae gastropod[[File:Physa acuta 001.JPGcenterthumb178x178pxExtant specimen]]
PleuroceridaeIndeterminate"Locality 4"ShellsIncertade Sedis
PomaceaP. sp."Locality 4"ShellsAn ampullariidae gastropod[[File:Pomacea canaliculata1.jpgcenterthumb178x178pxExtant specimen]]
PotamolithusP. fabiani"Locality 4"ShellsA tateid gastropod[[File:Potamolithus (10.3897-subtbiol.25.23778) Figure 4.jpgcenterthumb178x178pxExtant specimen]]
StenophysaS. sp."Locality 4"ShellsA physidae gastropod

Bivalves

GenusSpeciesLocalityMaterialNotesImages
*Chlamys***"C." salamanca"Locality 5"ShellsA Pectinidae bivalve[[File:Chlamys islandica 01.jpgcenterthumb225x225pxExtant specimen]]
CubitostreaC. ameghinoi"Locality 5"ShellsAn Ostreidae bivalve[[File:Cubitostrea digitata 01.jpgcenterthumb225x225pxExtant specimen]]
GryphaeostreaG. callophyla"Locality 5"ShellsA Gryphaeidae bivalve

Fishes

GenusSpeciesLocalityMaterialNotesImages
LamniformesIndeterminate"Locality 5"Isolated upper lateral toothSimilar to Cretalamna.
VidalamiinaeIndeterminate"Locality 2"Isolated toothAn amiid fish

Amphibians

GenusSpeciesLocalityMaterialNotesImages
AnuraIndeterminate"Locality 4"MPM 21518, isolated tibiofibula
CalyptocephalellaC. gayi"Locality 2"Incomplete left maxilla, incomplete right maxilla, incomplete right ilium, proximal right radioulna, left proximal end of tibiofibula and proximal half of urostyle[[File:Rana chilena.pngcenterthumb225x225pxExtant specimen]]
CalyptocephalellidaeIndeterminate"Locality 4"MPM 21519, distal end of right humerus with eroded distal margin
PipoideaIndeterminate"Locality 2"Distal end of right humerusResembles, yet is clearly different, from Xenopus.

Plesiosaurs

GenusSpeciesLocalityMaterialNotesImages
ElasmosauridaeIndeterminate"Locality 5"Incomplete cervical centrum and the lateral half of a very fragmentary dorsal vertebraMay belong to the Calafate Formation

Lepidosaurs

Mosasaurs

GenusSpeciesLocalityMaterialNotesImages
MosasauridaeIndeterminate"Locality 5"Seven teethIts presence suggest marine influences

Rhynchocephalian

GenusSpeciesLocalityMaterialNotesImages
last1=Agnolínfirst1=Federicolast2=Aranciaga Rolandofirst2=Maurolast3=Manabefirst3=Makotolast4=Tsuihijifirst4=Takanobulast5=Novasfirst5=Fernando E.date=2024-05-02title=Southernmost lepidosaur (Reptilia) assemblage from the Late Cretaceous of Patagoniaurl=https://www.tandfonline.com/doi/full/10.1080/08912963.2024.2341850journal=Historical Biologylanguage=envolume=37issue=4pages=888–902doi=10.1080/08912963.2024.2341850issn=0891-2963url-access=subscription}}N. finisterre"Monotreme site"Incomplete right maxilla preserving two completeand one partial teeth.A sphenodontid rhynchocephalian

Snakes

GenusSpeciesLocalityMaterialNotesImages
AlethinophidiaIndeterminate"Locality 4"MPM 21522 small vertebra
AnilioideaIndeterminate"Locality 2"MPM-PV-22847, possible mid-precloacal vertebra
ConiophiidaeIndeterminate"Monotreme site"Nearly complete mid-posterior precloacal vertebra; Incomplete posterior precloacal vertebraShares several features with Coniophis.
MadtsoiidaeIndeterminate"Monotreme site"; "Green Valley site"Incomplete mid-posterior precloacal vertebralacking most of the neural arch; Poorly preserved precloacal vertebra preserving the verteb-ral centrum
RionegrophisR. madtsoioides"Locality 2"A partial vertebra.A madtsoiid snake

Turtles

GenusSpeciesLocalityMaterialNotesImages
ChelidaeIndeterminate"Locality 2"MPM 21520, two peripheral and one costal platesLargest plates suggest a maximum carapace length of 40 cm.
HydromedusinaeIndeterminate"Locality 2, 4, 5"Left peripheral 3 or 4, bridge peripheral, fragment indet; bridge peripheral, right hypoplastron, buttress, plastral fragment and ?plastral fragment
MeiolaniformesIndeterminate"Locality 4"Distal end of a left humerus

Archosaurs

Crocodylomorphs

GenusSpeciesLocalityMaterialNotesImages
KostensuchusK. atroxEstancia La AnitaSkull, jaws, incomplete postcranial skeletonA large-bodied, broad-snouted peirosaurid[[File:Kostensuchus Simple Restoration.pngcenter200px]]

Ornithischians

GenusSpeciesLocalityMaterialNotesImages
last=Rozadillafirst=Sebastiánlast2=Agnolínfirst2=Federicolast3=Manabefirst3=Makotolast4=Tsuihijifirst4=Takanobulast5=Novasfirst5=Fernando E.date=2021title=Ornithischian remains from the Chorrillo Formation (Upper Cretaceous), southern Patagonia, Argentina, and their implications on ornithischian paleobiogeography in the Southern Hemispherejournal=Cretaceous Researchvolume=125article-number=104881doi=10.1016/j.cretres.2021.104881issn=0195-6671}}IndeterminateLa Anita FarmTwo incomplete caudal centra
author1=Fernando Novasauthor2=Federico Agnolinauthor3=Sebastián Rozadillaauthor4=Alexis Aranciaga-Rolandoauthor5=Federico Brissón-Eliauthor6=Matias Mottaauthor7=Mauricio Cerroniauthor8=Martín Ezcurraauthor9=Agustin Martinelliauthor10=Julia D'Angeloauthor11=Gerardo Álvarez-Herreraauthor12=Adriel Gentilauthor13=Sergio Boganauthor14=Nicolas Chimentoauthor15=Jordi García-Marsàauthor16=Gastón Lo Cocoauthor17=Sergio Miquelauthor18=Fatima Britoauthor19=Ezequiel Veraauthor20=Valeria Loinazeauthor21=Mariela Fernandezauthor22=Leonardo Salgadoyear=2019title=Paleontological discoveries in the Chorrillo Formation (upper Campanian-lower Maastrichtian, Upper Cretaceous), Santa Cruz Province, Patagonia, Argentinajournal=Revista del Museo Argentino de Ciencias Naturalesseries=Nueva Seriesvolume=21issue=2pages=217–293doi=10.22179/revmacn.21.655doi-access=freehdl=11336/122229hdl-access=free }}I. santacrucensis"Locality 5"Multiple specimens including teeth, several vertebrae, humerus, thigh bones, shoulder blades, and limbsAn elasmarian ornithopod[[File:Isasicursor skeletal.pngcenterthumb225x225pxIsasicursor skeletal]]
OrnithopodaIndeterminateLa Anita FarmProximal end of a femurA small sized ornithopod
ParankylosauriaIndeterminateLa Anita FarmFive fragmentary caudal vertebrae, incomplete ribs, two incomplete humeri, and nine osteoderms

Sauropods

GenusSpeciesLocalityMaterialNotesImages
FusioolithusF. sp."Locality 2"EggshellsFusioolithidae oogenus
LithostrotiaIndeterminateRío LeonaMACN 5205 (3 middle–posterior caudal centra)
NullotitanN. glaciarisEstancia Alta Vista; "Locality 1, 2, 4, 5"A partial skeleton consisting of a third cervical vertebra, tail vertebrae, a neck rib, ribs, a left shoulder blade, the ends of a right thigh bone, a right shin, a right calf bone, and a right ankle boneA colossosaurian titanosaur[[File:Nullotitan skeletal.pngframless200px]]
TitanosauriaIndeterminate"Locality 2, 4"Seven isolated teeth

Theropods

GenusSpeciesLocalityMaterialNotesImages
EnantiornithesIndeterminate"Locality 2, 4"Distal fragment of a pedal phalanx; Two isolated ungual pedal phalanges[[File:Maip macrothorax reconstruction.jpgthumbcenter200px[[Maip macrothorax]]]]
KookneK. yeutensis"Locality 2"An incomplete right coracoid, lacking the sternal end and the proximal end damagedAn ornithuran bird similar to Anseriformes
MaipM. macrothoraxLa Anita FarmAxis, several dorsal and caudal vertebrae, cervical and dorsal ribs, gastralia, a left coracoid, fragmentary scapula, partial right pubis, and partial metatarsalThe largest megaraptorid theropod of South America and possibly the largest member of this group
MegaraptoridaeIndeterminateLa Anita FarmIsolated teeth
NoasauridaeIndeterminate"Locality 4"Isolated right pedal phalanx IV-2?Similar to Velocisaurus and Vespersaurus
PrismatoolithidaeIndeterminate"Locality 4"EggshellsTheropod eggshells
UnenlagiidaeIndeterminate"Locality 4"An isolated dorsal vertebra, pedal ungual of digit II and phalanx III-2Similar to Neuquenraptor
YatenavisY. ieujensisEstancia La AnitaDistal right humerusAn enantiornithine bird

Mammals

GenusSpeciesLocalityMaterialNotesImages
MagallanodonM. baikashkenkeRío de Las Chinas Valley and La Anita FarmTentative first upper molariform, anterior end of a left dentary bearing part of its gliriform incisor, and upper mesial incisor from the right sideA ferugliotheriid gondwanathere
last1=Chimentofirst1=N. R.last2=Agnolínfirst2=F. L.last3=Moyano-Pazfirst3=D.last4=Manabefirst4=M.last5=Tsuihijifirst5=T.last6=Novasfirst6=F. E.year=2025title=New fossil mammal remains from the Chorrillo Formation (Maastrichtian, Upper Cretaceous), Santa Cruz Province, Argentinajournal=Journal of Vertebrate Paleontologyarticle-number=e2531263doi=10.1080/02724634.2025.2531263 }}"Locality 2"MPM-PV-23088, isolated right lower molariform; MPM-PV-23089, isolated right lower molariformA ferugliotheriid gondwanathere
MammaliaIndeterminate"Locality 4"Two isolated caudal vertebraeCharacters shared with mammals having long and gracile tails.
MeridiolestidaIndeterminate"Monotreme site"MPM-PV-21707, two fragments of a right dentarySmall sized taxon similar to Reigitherium.
MesungulatoideaIndeterminateLa Anita FarmMPM-PV-21708, fragment of left dentary with five alveoli corresponding with three teeth.Larger than any known Mesozoic meridiolestidan.
OrretheriumO. tzei"Monotreme site"MPM-PV-23514, left upper second molariform.A member of Meridiolestida belonging to the family Mesungulatidae.
PatagorhynchusP. pascuali"Monotreme site"Lower right molar attached to a fragment of the dentaryA ornithorhynchid monotreme[[File:Patagorhynchus holotype.pngcenterthumb189x189pxPatagorhynchus holotype]]
title=A large therian mammal from the Late Cretaceous of South Americafirst1=Nicolás R.last1=Chimentofirst2=Federico L.last2=Agnolínfirst3=Jordilast3=García-Marsàfirst4=Makotolast4=Manabefirst5=Takanobulast5=Tsuihijifirst6=Fernando E.last6=Novasdate=February 3, 2024journal=Scientific Reportsvolume=14issue=1page=2854doi=10.1038/s41598-024-53156-3pmid=38310138pmc=10838296bibcode=2024NatSR..14.2854C }}P. chainkoLa Anita FarmPartial left acetabulum and ischium; MPM-PV-23515, one fragment of thoracic vertebra, posterior half of one caudal vertebra and 38 indeterminate bones.A large therian[[File:Patagomaia chainko.webpcenterthumb225x225pxPatagomaia holotype]]

Flora

Algae

GenusSpeciesAffinity / FamilyImage
BotryococcusB. sp.Botryococcaceae
CymatiosphaeraC. sp.Zygnemataceae
LeiosphaeridiaL. sp.Zygnemataceae
OvoiditesO. parvusZygnemataceae
O. spriggiiZygnemataceae
SchizosporisS. reticulatusZygnemataceae (aff. Spirogyra)
SphaeropleaS. striatocristataSphaeropleaceae

Liverworts

GenusSpeciesAffinity / FamilyImage
AequitriraditesA. spinulosusMarchantiidae
StoverisporitesS. microverrucatusMarchantiidae
TaurocusporitesT. segmentatusMarchantiidae
TriporoletesT. reticulatusMarchantiidae
T. sp.Marchantiidae

Hornworts

GenusSpeciesAffinity / FamilyImage
ForaminisporisF. wonthaggiensisAnthocerotidae
F. sp.Anthocerotidae

Lycopods

GenusSpeciesAffinity / FamilyImage
CamarozonosporitesC. ambigensSelaginellales
CeratosporitesC. equalisSelaginellales
DensoisporitesD. velatusSelaginellales
FoveosporitesF. sp.Selaginellales
RetitriletesR. austroclavatiditesLycopodiales
R. sp. cf. R. douglasiiLycopodiales

Ferns

GenusSpeciesAffinity / FamilyImage
AzollaA. cf. andreisiiSalviniaceae
A. sp.Salviniaceae
BaculatisporitesB. comaumensisOsmundaceae
B. kachaikensisOsmundaceae
B. turbioensisOsmundaceae
cf. CladophlebisC. sp.Polypodiopsida
ClaviferaC. sp.Gleicheniaceae
CyatheaciditesC. annulatusDicksoniaceae (aff. Lophosoria)
CyathiditesC. australisCyatheaceae
C. minorCyatheaceae
C. rafaeliCyatheaceae
DictyophylliditesD. sp. cf. D. concavusDipteridaceae / Matoniaceae
GleicheniiditesG. sp.Gleicheniaceae
LeptolepiditesL. verrucatusBlechnaceae
OrnamentiferaO. sp.Gleicheniaceae
OsmundaciditesO. diaziiOsmundaceae
PeromonolitesP. vellosusBlechnaceae

Non-conifer Seed Plants

GenusSpeciesAffinity / FamilyImage
CycadopitesC. follicularisCycadales, Bennettitales, or Ginkgoales
EphedripitesE. ambiguousEphedrales
EquisetosporitesE. notensisEphedrales
E. sp. cf. E. dudarensisEphedrales
VitreisporitesV. signatusPteridosperms

Conifers

GenusSpeciesAffinity / FamilyImage
AraucariacitesA. australisAraucariaceae
CallialasporitesC. turbatusAraucariaceae
ClassopollisC. sp.Cheirolepidiaceae
CyclusphaeraC. psilataAraucariaceae
DacrycarpitesD. australiensisPodocarpaceae
DilwynitesD. tuberculatusAraucariaceae
GamerroitesG. psilasaccusPodocarpaceae
LygistepollenitesL. floriniiPodocarpaceae
MicrocachryiditesM. antarcticusPodocarpaceae (aff. Microcachrys)[[File:Microcachrys tetragona 112011800.jpgcenterthumb136x136px[[Microcachrys]]]]
PeripollenitesP. sp.Cupressaceae
PhyllocladiditesP. mawsoniiPodocarpaceae (aff. Lagarostrobos franklinii)[[File:Lagarostrobos franklinii 61295704.jpgcenterthumb181x181pxLagarostrobos]]
PodocarpiditesP. elegansPodocarpaceae
P. elipticusPodocarpaceae
P. exiguusPodocarpaceae
P. herbstiiPodocarpaceae
P. majorPodocarpaceae
P. marwikiiPodocarpaceae
P. otagoensisPodocarpaceae
P. sp. cf. P. langiiPodocarpaceae
P. sp. cf. P. microreticuloidataPodocarpaceae
P. torquatusPodocarpaceae
P. verrucosusPodocarpaceae
PodocarpoxylonP. duseniiPodocarpaceae
TrichotomosulcitesT. subgranulatusPodocarpaceae

Magnoliids

GenusSpeciesAffinity / FamilyImage
ClavatimonocolpitesC. sp.Magnoliids

Monocots

GenusSpeciesAffinity / FamilyImage
ArecipitesA. minutiscabratusArecaceae
A. sp. cf. A. subverrucatusArecaceae
Aff.Eolirion"Morphotype 5"Palmae
LongapertitesL. proxapertitoidesArecaceae
LiliaciditesL. lanceolatusLiliales (aff. Lilium)[[File:0 Lilium pyrenaicum - Samoëns 3.jpgcenterthumb203x203pxLilium]]
L. variegatusLiliales (aff. Lilium)
L. vermireticulatusLiliales
L. sp. cf. L. kaitangataensisLiliaceae/Amaryllidaceae
Monocotyledonean"Morphotype 6"Monocots
NupharN.? sp. 1Nymphaeaceae[[File:Nuphar lutea Krumpendorf 01.jpgcenterthumb204x204pxExtant specimen]]
N.? sp. 2Nymphaeaceae
PandaniiditesP. typicusAraceae
ProxapertitesP. operculatusAraceae
SpinozonocolpitesS. hialinusArecaceae

Eudicots

GenusSpeciesAffinity / FamilyImage
BeaupreaB. elegansiformisProteales[[File:Beauprea filipes.jpgcenterthumb200x200pxBeauprea drawing]]
Aff. Brachychiton"Morphotype 4"Malvaceae[[File:Brachychiton acerifolius Royal National Park.JPGcenterthumb181x181pxExtant specimen]]
ButtiniaB. andreeviiBuxales
ClavamonocolpitesC. sp.?Chloranthaceae
Coccoloba?"Morphotype 1"Polygonaceae[[File:Uva de playa (Coccoloba uvifera) unsharp.jpgcenterthumb183x183pxExtant specimen]]
ConstantinisporisC. jacquei?Chloranthaceae
DicotetraditesD. sp.Ericales
DiporitesD. sp.?Chloranthaceae
DoyleipollenitesD. sp.?Chloranthaceae
EricipitesE. scabratusEricales
Eudicots"Morphotype 3"Eudicots
MyrtaceiditesM. sp. cf. M. mesonesusMyrtales
NothofagiditesN. saraensisNothofagaceae (cf. Nothofagus)[[File:Scheinbuchenwald (Nothofagus) Torres del Paine Nationalpark.jpgcenterthumb136x136px[[Nothofagus]]]]
NyssapollenitesN. sp.?Chloranthaceae
PeninsulapollisP. gilliiProteaceae (cf. Beauprea?)
P. truswelliaeProteaceae (cf. Beauprea?)
PolycolpitesP. sp.Pedaliaceae (?cf. Josephinia)
ProteaciditesP. parvusProteaceae (cf. Bellendena montana)[[File:Bellendena montana - Walls of Jerusalem.jpgcenterthumb136x136px[[Bellendena montanaBellendena]]]]
P. subscabratusProteaceae
P. sp. cf. P. tenuiexinusProteaceae
P. sp.Proteaceae
PsilatricolporitesP. sp. cf. P. complanatiusTetrameristaceae
P. sp.Tetrameristaceae
RetidiporitesR. camachoiProteaceae
RhoipitesR. baculatusEricales
R. minusculusEricales
R. sp. cf. R. alveolatusEricales
R. sp. 1Ericales
R. sp. 2Ericales
RouseaR. patagonicaEricales
R. sp. cf. R. minusculaEricales
R. sp.Ericales
SyncolporitesS. sp.Ericales
TetracolpitesT. sp.?Chloranthaceae
TriatriopollenitesT. lateflexusProteaceae
TricesticillusT. americanus?Chloranthaceae
TricolpitesT. reticulatusGunneraceae (Gunnera macrophylla)[[File:Gunnera leaves, Coleton Fishacre - geograph.org.uk - 2427461.jpgcenterthumb171x171px[[Gunnera macrophyllaGunnera]]]]
T. sp. cf. T. brevicolpusGunneraceae
T. sp. cf. T. inargutusGunneraceae
T. sp. cf. T. lumbrerensisGunneraceae
T. sp. 1Ranunculaceae (cf. Clematis)[[File:Clematis Montana 02.jpgcenterthumb181x181px[[Clematis]]]]
TricolporitesT. sp. 2Ericales
T. sp. 3Ericales
TricolporoiditesT. sp. cf. T. eximiusEricales
TrioritesT. sp. cf. T. minusculus?Chloranthaceae
TubulifloriditesT. lillieiEricales
VolkheimeritesV. labyrinthus?Chloranthaceae

References

Bibliography

References

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  2. Marenssi et al., 2003, p.406
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